Team Dominator risks everything to warn people in the path of a violent storm, but internal conflicts causes driver Joel to leave, complicating Team Dominator's operations. Still gun shy after witnessing devastation this season, Sean Casey starts to lack the passion of chasing storms, until a visit from his wife and a successful chase reignites Team TIV's fire. Team TWISTEX tries to brave the elements by chasing a tornado through a hail storm, but after their vehicles suffer damage, Tim has to swallow his pride and abandon the chase.
In the wake of the deadly and very large tornado outbreak (April 27, 2011), all three chase teams take time to help with the recovery effort. Team Tiv is in Smithville, Mississippi and Team Dominator and Team Twistex are in Tuscaloosa, Alabama. Motivated by the death and destruction they've witnessed, all three teams are rededicated to improving warning times and doing vital research on these deadly storms. Team Dominator fields a new, improved vehicle: the Dominator 2 and tracks a storm right toward one member's childhood home. Team TWISTEX outfits their truck with a brand new probe and a mobile Doppler radar, but mechanical issues threaten to sideline them again. Team TIV adds a mobile tornado siren and chases toward the heart of a twister to warn people while shooting 3D twister footage.
After failing to find a twister on two consecutive chases, Reed Timmer and the Dominator team are desperate to chase one down. They're even desperate enough to pursue a powerful storm into an urban area after dark, one of the most dangerous possible chasing scenarios. Team TWISTEX's truck threatens to keep them on the sidelines on a promising tornado day. While team Dominator pursues an intercept, the TIV team is hoping to finally shoot their first ever 3D tornado footage... but tension in the vehicle has everyone on edge, and the crew is finding out that the only thing worse than having a chase ruined by the slick, Texas mud is to be helpless in the path of a tornado.