Niles' persistence finally convinces Daphne's father to come to Seattle to reconcile with his wife. Daphne is disappointed that Niles didn't consult her first, but is on board with the plan.
Guilt feelings over a sexual liaison with Roz plague Frasier, who heads out to her family reunion in Wisconsin to make amends.
Frasier's show has branched out to Spokane, and he goes to the station in Spokane to meet the press. He quickly finds out that the press and his callers refuse to warm up to him.
Frasier returns to Boston to give a speech, and Niles, Daphne and Martin tag along to see the city. Once there, Frasier runs into Cliff Clavin.
To quell a feud between Frasier and an upstairs antagonist, Martin and the neighbor's mother feign a romance.
While Frasier and Niles uncover a human skull under the floor of their childhood home and let their imagination get the better of them, Roz's daughter's hamster has died and she is uncertain how to break it to her.
A-N-X-I-E-T-Y seizes Frasier at the National Spelling Bee, where his son Frederick is a leading competitor.
Niles sets Frasier up with Lisa, a patient who owns a bookstore, but they never run into each other. Roz sets Frasier up with Suzanne, an artist, who ends up hating Frasier. Martin sets Frasier up with Kris, an attractive girl from his building, who seems to know every man at the bar. But Frasier later runs into and hits it off with Lisa.
Frasier is unimpressed when his scheming, scamming ex-brother-in-law (Michael Keaton) rolls in for a visit -- now in a wheelchair -- and claims to be a reformed, God-fearing man who needs money for a revival ministry.
Niles plans on proposing to Daphne, so he prepares his speech; buys a ring; and hires a choir, a string quartet, a trumpet player and Wolfgang Puck. But Daphne's mother is still staying with them, and on the night Niles plans to propose, Daphne begins to feel ill.
Kenny tells Frasier and Roz that some recent research into KACL listeners shows that Frasier's average listener is an older gentleman who keeps the radio on for company. He suggests Frasier reach out to the youth.
Daphne and Niles continue to try and hide the fact that they have moved in with each other from Mrs. Moon. She makes such a mess of things that the only way she will leave is if Simon takes care of her, but Simon has left for California. Meanwhile, Frasier's neighbor Cam drapes the American flag over his balcony, and Frasier cannot complain because it would seem unpatriotic.
When Daphne and Niles decide to move in with each other, Daphne's mother and Simon arrive on a surprise visit. Daphne and Niles decide to put off moving in together until her visiting relatives are gone. Meanwhile, Frasier feuds with Cam Winston, his neighbor, over where Cam parks his large vehicle.
After Frasier witnesses Martin get chewed out by his boss, Frasier feels obligated to say something on his behalf. However, it only serves to make things worse.
Frasier gets an annoying new agent in the middle of a competition between him and a rival radio broadcaster.
Frasier and Niles compete to see some rare wine, owned by a rich relative of Kirby's girlfriend. Meanwhile, Martin must wear a blood pressure monitor.
Frasier has just completed his 2000th radio show, but discovers that one of the tapes of his collection of his broadcasts is missing. His quest to replace it borders on obsession, sending him spiralling into depression.
An accident causes a mess on Frasier's carpet and the destruction of Martin's chair. Meanwhile, Niles gives piano lessons to talented Roz and not-so-talented Daphne.
Frasier holds a 'Dress as Your Hero' Halloween party, but nobody can tell who he is. Meanwhile, Niles dresses as Martin, Daphne as Elton John, Martin as Joe DiMaggio and Roz as Wonder Woman for the party.
Frasier hosts a party for the staff at the radio station to improve his image. Meanwhile, Roz dates a garbageman and Niles recalls his childhood memories with Daphne.
Martin goes back to work as a security guard and reflects on his last day as a cop, when he was shot and had to retire.
Daphne stumbles across a file of one of Niles' patients, who is in love with him via "transference." Daphne's paranoia sends her to extreme lengths to learn anything she can about the woman.
Troubled by his recent relationship woes, Frasier goes to the country for peace and quiet but ends up facing the ghosts of failed relationships past.
Frasier continues to be conflicted about whether he truly wants to be with Lana instead of Claire. He consults his family, but a chance encounter at the airport steers him toward making a decision. Unfortunately, he may be too late.
