Ralf Little heads to Orkney to investigate his grandfather Arthur’s role in one of the most important battles of the Second World War. He also finds an international footballer in the family.

Motherland star Anna Maxwell Martin travels to the small town in Northern Ireland where her father grew up, discovering the dark story of her grandfather's childhood.

Matt Lucas learns what his late grandmother went through as a Jew in Nazi Germany in the 1930s, and what happened to the family she left behind. He also uncovers a connection to Anne Frank.

Richard Osman delves into his Brighton roots and is astonished to learn that one of his ancestors turned amateur sleuth and ended up entangled in one of Brighton’s most notorious murder trials.

Sue Perkins delves into the German branch of her family, uncovering a harrowing tale of refugees fleeing back and forth across east European borders during the Second World War.